Obligate & Facultative Seeders

My elegant crossing Tule Creek. Photo by Josh Smith.

Last weekend, I hiked in Trinity County along a low-elevation, fire-prone section of the Bigfoot Trail between Highway 3 and Hayfork and was able to witness obligate & facultative seeding in action.

From Field Guide to Manzanitas, Backcountry Press, 2012

One-third of manzanita species are facultative seeders. These are species that regenerate post-fire by both seed and burl resprouting. The remainder are obligate seeders that lose their entire adult population in a fire and depend on a seed bank for regeneration. Obligate seeding is the current model in manzanita evolution.

To understand why, consider the climatic dynamics over thousands, or tens of thousands of years or more. In the case of the resprouting species, particular individuals can live for centuries, resprouting over and over, cloning new individuals as the burls expand with each fire cycle. But in that population, the rate of genetic change is limited, because most individuals live a long time by way of asexual reproduction. This suggests that populations may be unable to respond to rapid climatic changes that might occur in only hundreds of years. The obligate seeders, on the other hand, lose all adults in stand-replacing fires and new post-fire generations have to establish from more genetically diverse seeds. Those populations consequently have greater flexibility to shift and adjust as circumstances require; traits that might have been rare and less important in older generations can emerge through natural selection and become critical in the newer generations within the lifetime of resprouting manzanitas.

Much of this area burned in the summer of 2015. While evidence of the fires were everywhere, there are many signs of the next generation of plants returning to the landscape. This was particularly true on some of the south-facing slopes above Philpot Campground where two species of manzanitas were exploring different reproductive regimes– both obligate & facultative seeding.

Arctostaphylos manzanita is a  facultatative seeder. It can resprout by seed but also from dormant buds. When present, these structures are often prominent and are called burls (also ligno tubers)–seen in the middle of the sprouting leaves.
Arctostaphylos canescens is an obligate seeder and does not have a burl. Instead, genetically unique seedlings resprout from a well-stocked seed bank.

Nature Notes:

A large burned stand of the obligate-seeding hoary manzanita, above Philpot Campground on the Shasta-Trinity National Forest.

Montgomery Woods State Natural Reserve

Albino redwood.
Albino redwood.

Hot springs and tall trees, all hidden in a remote area of Mendocino County. This was the family destination for a pre-Thanksgiving celebration. Most or our time was spent at Orr Hotsprings but an day-hiking excursion treated us with a fabulous adventure in an isolated redwood grove at Montgomery Woods State Natural Reserve.

The hike was punctuated with firsts: my son’s first multi-mile hike and my my first albino redwood.  The park previously held the tallest recorded redwood — at 367.5-foot it was once declared the world’s tallest. While the Montgomery tree is still alive, the new record  is in Humboldt County. Other exciting finds in the park included an excellent expanse of giant chain ferns in a lowland basin along the trail. Read more about Montgomery Woods from Save the Redwoods League.

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Arctostaphylos nummularia

Arctostaphylos nummularia subsp. mendocinoensis

The pygmy manzanita (Arctostaphylos nummularia) is a species endemic to Mendocino County, California  where it is known from its occurrence in the pygmy forests along the coastline. I think this might just be the perfect northern coastal California shrub for a native plant garden. It has a perfectly rounded form, beautiful small leaves, and subtly hairy stems.

Arctostaphylos nummularia
Arctostaphylos nummularia subsp. mendocinoensis by Allison Poklemba

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Forest Pathogens

Select Forest Pathogens of California’s Klamath Mountains

Forest Pathogens often go unnoticed while exploring, but offer an exceptional window into the intricacies of forest ecology when better understood. I created the free document linked below in 2011 while in Grad School at Humboldt State. Forest Pathology was one of the more interesting classes I took while turning Conifer Country into my thesis for a Master’s Degree in Biology. Most of the information for this document was taken from Terry Henkel’s lecture notes as well as from internet and book sources–all cited within the document. I was recently reminded of this creation because of the October weather that has dropped unprecedented amounts of rain and nurtured fungal growth across Northwest California.

Fomitopsis pinicola
Fomitopsis pinicola

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Bear Peak Botanical Area

Citizen Science in the Siskiyous

img_4865I recently started a citizen science project with 5 classes of high school biology students from Fortuna, California. The plan is to combine their observation skills with the technology offered by iNaturalist. Each month they will visit Rohner Park and record data on a chosen spot in the forest–looking for plants and animals as well as  changes in canopy and ground cover. As they become more proficient in species ID, students will also upload observations to our iNatural Project ultimately creating a field guide to their local forest. We all know how much I like field guides…

My plan, over future visits to wilderness areas, is to start similar citizen science projects.  The first attempt at this wide-ranging project began this week on a visit to the Bear Peak Botanical Area on the Klamath National Forest. I originally wrote about this area in my book Conifer Country because it is unique in many ways, including the populations of yellow-cedar found  here. This species in common further north, but quite rare in California.

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My Favorite Fall Hikes

Quaking aspens (Populus tremuloides) in the Blackrock-High Rock Desert.
Quaking aspens (Populus tremuloides) in the Blackrock-High Rock Desert.

Fall is coming.

For me it is a time of rejuvenation and also my favorite season. With the shift of the California Current, rains begin to fall in California after a summer of drought. The high country along the Pacific Slope finds snow returning. While we retreat inside our homes, native plants and animals must adjust to the changes. Some birds migrate, mammals might hibernate, and some plants shed their leaves and “hibernate” for winter in their own way.

What follows is a journey across the Pacific Slope to four favorite fall hikes–excellent for colorful foliage. I’ve also included some of the plant species that will be found.
  1. Siskiyou Wilderness
  2. Pasayten Wilderness
  3. San Gabriel River National Monument
  4. Blackrock-High Rock Desert National Conservation Area

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Mount Linn Revisited

Northern California’s Coast Range

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A view from Mount Linn toward the Pacific Ocean and the distant King Range.

Mount Linn–also called South Yolla Bolly Mountain–is the highest point in the Coast Range of northern California. It is located to the west of Corning but the area might as well be a world away from the population centers of the state; it is rarely noticed by travelers as they drive Interstate 5. Once off the interstate, scenic forest service roads still take nearly 2 hours to wind to the trailhead. I revisited this fine mountain in July of 2016 to set up a photo-monitoring plot along the Bigfoot Trail and took the time to also map the vegetation on the mountain–particularly the grove of foxtail pines near the summit. This is one of the smallest (12 acres) and most isolated groves for the entire species and one that I am very much concerned about due to climate change. Shasta firs are encroaching upon the trees as snowpack declines and temperatures warm. I was happy to see the trees doing well and many young foxtails sprouting up–just not as many as there are young firs.

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Middle Tennessee Cedar Glades

Map adapted from William J. Wolfe, “Hydrology and Tree-Distribution Patterns of Karst Wetlands at Arnold engineering Development Center,” USGS

Edaphic islands have long-fascinated me–especially having grown to understand the serpentine barrens of the Klamath Mountains. So when we found ourselves visiting the Central Basin Region of middle Tennessee I discovered, through the help of the Tennessee Native Plant Society, that this area is botanically special for its limestone prairies, often called glades. While numerous state designated natural areas were recommended, I chose to learn about the Middle Tennessee Cedar Glades at Flat Rock Glades and Barrens State Natural Area.

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Choosing a Hike

How do you go about choosing a hike?

I have used various approaches which always involve careful map study, perusing the pages of hiking guides, and most importantly for me—studying field guides. As I get older, choosing a hiking destination is becoming more critical, with so much to see and even more to learn.

Hiking in the King Range.
King Range hiking.

Over time, I have gone about choosing a hike based more as a destination for discovery before any other factor. I think I first caught the hiking-for-natural-discovery bug while selecting a backpacking route exclusively to see condors in the Sespe Wilderness of southern California. When I moved to Humboldt in 2002, I graduated from bird destinations to plant exploring as I began searching out rare and unusual conifer species in our local mountains. This regular wilderness sideline blossomed into a Master’s Degree from Humboldt State University when I published my first book Conifer Country: A natural history and hiking guide to the conifers of northwest California in 2012. For 10 years I hiked to find and understand trees. These trees, and the places they grow, helped me develop a deeper passion for place and an understanding of the unique natural history of northwest California.

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Record Monterey pine (Pinus radiata)

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Circumference of Humboldt County’s record Pinus radiata

I have known about this Monterey pine (Pinus radiata) for years.

Pines in a redwood forest? Yep!
Pines in a redwood forest? Yep!

In 2004, I first took my 7th graders from Fortuna Middle School to make observations in their science journals in the forest surrounding this beauty. I always knew it was big, but did not know it could be the largest of its species.

Champion Pinus radiata
Champion Pinus radiata

The history of the forest at Rohner Park is not well documented, but as luck would have it I found a few answers while measuring the tree. With laser rangefinder in hand, an old-timer from Fortuna was coincidentally walking past me and asked what I was doing. His understanding was that Boy Scouts had planted a handful of pines in the area after the old-growth redwood forest was logged by cross-cut saw–in the years just before the invention of the chain saw in the 1920s. That would mean that these trees were, most likely, planted between 1900-1920. This makes the Monterey pines here, and the mature second-growth redwood forest, nearly 100 years old.

It is an impressive tree, competing with the forest giants of the North Coast like redwoods, Douglas-firs, and grand firs. Conditions must be right for this pine to survive among these other shade-tolerant trees. Pines, remember, are usually not shade-tolerant. That being said, if this tree was planted before the redwoods re-grew after logging in the early 1920s, then it got a head start and grew tall, in a race for light, against the other species in the park. Amazing stuff, to see a 160′ pine eking out an existence in the rainforest!

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San Gabriel Mountains Presentation

Plant Exploring in the San Gabriel Mountains National Monument

Wednesday, May 11, 2016 @7:30 p.m. at the Arcata Masonic Lodge

From the California Native Plant Society North Coast Chapter:

Explorer, writer, and educator Michael Kauffmann will lead us on a journey into the Transverse Ranges of southern California to explore the world of what John Muir called the steepest mountains in which he ever hiked. Michael’s explorations began because of a Bigcone Douglas-fir mapping and monitoring project he is leading in conjunction with California Native Plant Society, but these studies have allowed him to make more discoveries–from one of the world’s largest oaks to the most isolated grove of Sierra junipers in the world. Michael will take us on a photographic journey from the mountain tops to the river canyons across one of the nation’s newest national monuments.

San Gabriel Mountains

Klamath Mountain Snowpack

Pondering the 2016 hiking season…

I have tracked how Klamath Mountain snowpack is correlated with the beginning of summer hiking season since 2003. In 2009 I started the Bigfoot Trail on the summer solstice, and it worked out perfectly—I found the perfect mix of snow, open trail, and manageable river crossings. 2010 was a late snow year and June was only open for hiking in the Southern Siskiyous. In 2011 I began a section hike from the Trinity Alps to the Siskiyous on June 29th in a snowstorm and we trekked across snow for nearly two weeks while in the Trinity Alps and Marble Mountains. Our time also included numerous, often stressful, fords of raging rivers in the low-country. What follows is an analysis of the El Niño winter we are emerging from and what snowpack trends seem to be saying about the 2016 hiking season.

2011-Alps
2011 section hike — crossing Stuarts Fork in the Trinity Alps and snowfields in the Marble Mountains

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California’s Botanical Landscapes

A PICTORIAL VIEW OF THE STATE’S VEGETATION

By Michael G. Barbour, Julie M. Evens, Todd Keeler-Wolf, John O. Sawyer

I am very fortunate to have been a part of the book project now in print titled CALIFORNIA’S BOTANICAL LANDSCAPES: A PICTORIAL VIEW OF THE STATE’S VEGETATION (CNPS Press 2016, $39.95). Over the past 5+ years this book has evolved through volunteer efforts as a service to those passionate about California’s flora. The book is dedicated to Humboldt State University botany professor and North Coast CNPS founder John O. Sawyer. I am honored to have contributed time to the project with photographs for many of the ecoregions as well as coauthoring the Klamath Mountain chapter with Sawyer. This new book surveys our state’s native vegetation with photos and text exploring each of 14 ecoregions across the state. It includes a wide array of photographs of broad-scale vegetation patterns paired with in-depth, interpretive descriptions written by California’s top plant ecologists. 

California's Botanical Landscapes
California’s Botanical Landscapes

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Owen Cheatham Redwood Grove

Or…Why Star Wars fans should love conifers–especially redwoods

Return of the Jedi had a major influence on my experiences as a youth. I wanted to be able to use the force like Luke Skywalker, as do many children again in 2016. The Star Wars phenomenon has trickled down to our 3 year-old son from his Kindergarten classmates at Montessori as well, so we decided to seize the day and go for a hike that combined the legend of the force with the power of the redwoods, and explore where science fiction meets natural history.

redwood needles and cones

Redwoods might just be the grandest species in the plant world—from top to bottom they are the epitome of grandeur. The tallest redwood is 379’ (and still growing), while one of the most massive redwoods is a mere 320’ but has a basal diameter of nearly twenty-six feet. Those are some impressive beings! The largest redwoods grow in northwest California, where they favor mountain slopes and river terraces close to the ocean while also being situated in the summer fog belt. Across the coastal landscape redwoods domineer a highly specific range—when soil, water, and exposure are optimal. The Owen Cheatham Redwood Grove is a great place to visit these giants.

In search of redwoods and film history in the Owen Cheatham grove.
In search of redwoods and film history in the Owen Cheatham Grove.

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San Gabriel Mountains – A Photo Tour

Celebrating the Bigcone Douglas-fir of the San Gabriel Mountains

After nearly a month of travel through one of the gems of Southern California, I’ve had enough time to reflect on all aspects of the journey–including the wonders of the wilderness and forest, as well as the state of the region. The San Gabriel Mountains remain wild, in large part, because of isolation due to slope. John Muir called them the steepest mountains he ever hiked in! It seems to me that the forests here are doing as well as they are, while our climate is rapidly changing, because of this isolation in slope. Forest pathogens travel much slower through heterogeneous landscapes with mixed stands of trees. Many of the Bigcone Douglas-fir stands we visited were in isolated on slopes of greater than 50º. Isolation sculpts the ecology of the mountain’s biota  in many ways, and makes life for Pseudostuga both easier and more difficult. That balance defines the ecological amplitude of many of the species on the California Floristic Province.

I’m still working on the full report for our findings, but in general it can be said that within region 1 of the map below, Bigcone Douglas-fir are not doing well. Elsewhere, they seem to be doing fine at the moment–especially if the species has been able to avoid high intensity fires. Reproduction is occurring at variable rates but they seem to like disturbed areas, like landslides, which the San Gabriels have no problem offering.
map-PSMA

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Manzanitas of the San Gabriel Mountains

Botanical Wonders of the San Gabriel Mountains National Monument.

Manzanitas are most commonly found across the Angeles National Forest on south-facing slopes where they are restricted to various mineral soils (most often granites). A. glandulosa appears in the front range from ~2,000-5,000 feet, A. patula in the higher elevations above ~7,000, A. parryana on the north slopes toward the Mojave from ~5,000-7,500, and A. glauca is common across the range but mostly on the north slopes adjacent to the Mojave from ~4,000-6,000. I never found Arctostaphylos pungens but it is within the range according to various sources.

What follows is a photographic journey through the San Gabriel Mountains to enjoy the spectacular places manzanitas grow.

glandulosa
Arctostaphylos glandulosa ssp. gabrielensis decorate the beginning of the Devil’s Canyon trail near Chilao Flat.

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Record Bigcone Douglas-firs

Exploring the San Gabriel Mountains National Monument

In my search to understand Bigcone Douglas-fir (Pseudotsuga macrocarpa) within the Angeles National Forest, I found myself in secret canyons and along steep hillsides that few people have ever explored. I thus took it upon myself to document more than just our selected data plots for Bigcone. In previous posts, I’ve shared some of the large angiosperms I’ve run across. Here is the documentation of the second largest Pseudotsuga macrocarpa yet measured — a record Bigcone Douglas-fir.

This Bigcone was found on the edge of a wash called Holcomb Canyon within the Devil’s Punchbowl Natural Area. The tree is nearly as big around as the record specimen in Baldy Village but just not as tall. I have heard a rumor  that the Baldy tree lost some of its crown — so I wonder if the numbers I have for this tree are still correct. Regardless, the tree in the Punchbowl is much more dramatic, being that it is within the Pleasant View Ridge Wilderness and not in the middle of town!

Devils_PSMA

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Alnus rhombifolia

Alnus-Rhombifoilia-Kauffmann-blog2
Record white alder within the Pleasant View Ridge Wilderness.

I need to start by clarifying something: in last post about a giant Canyon Oak, I mentioned that I was not a big tree hunter. This post, a few days later, is about a big tree. I think I have become a big tree hunter…

Every day for the better part of the last two weeks, I have been walking through the San Gabriel Mountains getting to know Bigcone Douglas-fir as part of a project with the Angeles National Forest and the California Native Plant Society. I’ll post more about that in the near future. For now, lets look at the white alder (Alnus rhombifolia) I measured in the Pleasant View Ridge Wilderness within the San Gabriel Mountain National Monument while working on that project. Stats on the past record tree can be found HERE.

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Canyon Live Oaks of Southern California

The Transverse Ranges hold some of the largest oaks in North America

I fell in love with this Canyon Live Oak (Quercus chrysolepis) in 2002 while training to prepare myself to hike the Continental Divide Trail. One fateful night I inadvertently camped near it along a stream called Prairie Gulch. The tree has lived in my dreams ever since. In November 2015, the opportunity to return, revisit and measure the oak was offered  while doing a botany project in the San Gabriel Mountains National Monument.

The San Gabriel Oak
The San Gabriel Oak

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