In mid-September I had the opportunity to assist in guiding an EcoFlight across the North Coast and Klamath Mountains. I was invited by the California Wilderness Coalition to point out landmarks and important areas to Congressman Huffman’s staff and local media outlets as they relate to the Northwest California Wilderness, Recreation, and Working Forests Act. The following is a photo tour from the flight that highlights parts of the bill. If so inspired, please follow the links at the end to get involved!
Coastal lowland forests of the Klamath Mountains. From the future book, The Klamath Mountains: A Natural History.
The coastal lowland forests receive the most annual rainfall of anywhere else in the Klamath Mountains. From the north in Oregon the Coquille, Rogue, and Illinois rivers drain southwest Oregon and the Smith River country is California’s only temperate rainforest. The rich soils, temperate year-round growing season, and high rainfall (often over 100 inches) nurture these impressive forests.
Klamath coastal lowland forests are typified by dense, closed canopies on well-developed soils (sometimes serpentines). The dominant tree species have high colonizing abilities, long life, and a wide ecological tolerances to environmental conditions, including dry summers with occasional persistent fog. These forest thrive below the snow belt on the extreme western slopes of the Siskiyou Mountains. Species like Douglas-fir (Pseudotsuga menziesii) and Port Orford-cedar (Chamaecyparis lawsoniana) secure the highest place in the canopy, often approaching 300’. A second canopy forms one hundred feet below and includes species like tanoak (Notholithocarpus densiflorus) and madrone (Arbutus menziesii) where they share an understory with coastal specialists like salal (Gaultheria shallon) and evergreen huckleberry (Vaccinium ovatum). With the high volumes of rain that falls each winter, banks of coastal rivers and streams are frequently disturbed and then repeatedly pioneered by red alder (Alnus rubra).
Botanizing the South Fork National Recreation Trail – Trinity County
Hiking Hell Gate
Hell Gate highlights the lowland interior forest of the southern Klamath Mountains along the Trinity River on the Shasta-Trinity National Forest. The Trinity is surely one of the most spectacular rivers in the state and this hike is along a roadless stretch in the upper reaches of the south fork. The old-growth Douglas-fir/mixed evergreen forest is top-notch, especially along the north-facing sections of trail. Oak woodlands are on benches above the river, with Oregon white oak mixing with picturesque grasslands–though the lack of fire is allowing extensive conifer encroachment. Some of the largest Pacific yew I have ever seen can be found along the trail as well.
The area is dotted with private in-holdings and these beautiful old homes appear at random intervals along the trail. The trail is open to biking and—for these home-owners—OHVs. Swimming opportunities abound and solitude can be easily found, especially in the winter (there are bridges on all major stream crossings). The hike, as written and drawn in Conifer Country, is 7 miles to Smokey Creek but the National Recreation Trail extends the length of this roadless stretch of river—for a total of 15 miles from Hells Gate to Wild Mad Road. The upper reaches of this trail is part of the Bigfoot Trail.
I am highlighting three species below that I found on our hike. One that is a regional endemic, one that reaches it coastal range extension here, and a third that is found across the northern hemisphere. These selections highlight the regional diversity with a biogeographical perspective.
From Wildflowers of the Trinity Alps. Photo by Ken DeCamp.
Green plants are considered autotrophs because they photosynthesize—making sugar from water and carbon dioxide. The world of heterotrophic plants is complicated but all have moved away from total energy production from photosynthesis toward obtaining organic carbon either directly from other living beings or through a parasitic relationship with a fungus. Heterotrophic plants include directly parasitic and mycotrophic forms. The conifer forests of the western United States nurture an exceptional diversity of heterotrophic plants and the Klamath Mountains may have the most heterotrophic plants in the western United States!
My elegant crossing Tule Creek. Photo by Josh Smith.
Last weekend, I hiked in Trinity County along a low-elevation, fire-prone section of the Bigfoot Trail between Highway 3 and Hayfork and was able to witness obligate & facultative seeding in action.
One-third of manzanita species are facultative seeders. These are species that regenerate post-fire by both seed and burl resprouting. The remainder are obligate seeders that lose their entire adult population in a fire and depend on a seed bank for regeneration. Obligate seeding is the current model in manzanita evolution.
To understand why, consider the climatic dynamics over thousands, or tens of thousands of years or more. In the case of the resprouting species, particular individuals can live for centuries, resprouting over and over, cloning new individuals as the burls expand with each fire cycle. But in that population, the rate of genetic change is limited, because most individuals live a long time by way of asexual reproduction. This suggests that populations may be unable to respond to rapid climatic changes that might occur in only hundreds of years. The obligate seeders, on the other hand, lose all adults in stand-replacing fires and new post-fire generations have to establish from more genetically diverse seeds. Those populations consequently have greater flexibility to shift and adjust as circumstances require; traits that might have been rare and less important in older generations can emerge through natural selection and become critical in the newer generations within the lifetime of resprouting manzanitas.
Much of this area burned in the summer of 2015. While evidence of the fires were everywhere, there are many signs of the next generation of plants returning to the landscape. This was particularly true on some of the south-facing slopes above Philpot Campground where two species of manzanitas were exploring different reproductive regimes– both obligate & facultative seeding.
Arctostaphylos manzanita is a facultatative seeder. It can resprout by seed but also from dormant buds. When present, these structures are often prominent and are called burls (also ligno tubers)–seen in the middle of the sprouting leaves.
Arctostaphylos canescens is an obligate seeder and does not have a burl. Instead, genetically unique seedlings resprout from a well-stocked seed bank.
